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Modern weights and morphometric measurements of the cheetah (Acinonyx Jubatus)

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Arnoldia Rhodesia 1967


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The Mammals of the Southern African Subregion


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Cheetah skull measurements and wieghts from Comparative bite forces and canine bending strength in feline and sabretooth felids: implications for predatory ecology 


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( This post was last modified: 05-31-2021, 02:38 AM by Acinonyx sp. )




Cheetah gets weighed and measured.

Weight of the specimen


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Same specimen being measured


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Asiatic cheetahs

A review of ecology and conservation status of Asiatic cheetah in Iran
In Iran, we collected morphological data from 18 cheetahs from the subspecies range in Iran as well as literature (Hunter et al. 2007; Supporting Online Material SOM Table T1). Adult Males weight range from 25 to 38 kg while females vary between 23 and 35 kg, resulting in a smaller body size of Asiatic cheetahs than the Africans (SOM T1).

Conserving the Asiatic Cheetah in Iran: Launching the first radio-telemetry study

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^Morphological data for 2 male cheetahs.
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( This post was last modified: 05-31-2021, 06:51 AM by Acinonyx sp. )

Human-Cheetah size comparison made by me 


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Cheetah measurements and body mass from Wild Cats of the World


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( This post was last modified: 06-13-2021, 07:30 AM by Acinonyx sp. )

Body mass of the Cheetah (Acinonyx Jubatus) in wild,captive and semi-captive populations


Namibia
Etosha National Park
Cheetahs in Etosha national park,Namibia weigh 43 kg(n=306) for males and 36kg (n=154) for females.The biggest male cheetah in Etosha national park weighed 59 kg and the smallest male from the same place weighed 39 kg.The biggest female in Etosha national park weighed 45kg and the smallest female from the same place weighed 30 kg. For both genders cheetahs average 41kg (n=460) in Etosha national park.(1)


Namibian Farmlands
In namibian farmlands male cheetahs weigh 46 kg(n=108) and female cheetahs in the same place weigh 37 kg (n=43).As a species cheetahs weigh 44kg(n=142) in Namibian farmlands.The biggest male cheetah ever recorded in Namibian Farmlands weighed 64 kg and the smallest male cheetah ever recorded in the same place weighed 31kg.The smallest female cheetah ever recorded in the namibian farmlands weighed 26 kg and the biggest cheetah ever recorded from the same place weighed 51 kg.(4)(16)(18) 


Africat (Namibia)


Africat gives 46 kg average for male cheetahs (n=89 ) and 39 kg average for females (n=68).For both genders cheetahs averaged 43 kg (n=157) in this sample.The biggest male recorded by Africat weighed 62 kg.The biggest female recorded by Africat weighed 46 kg.(3)


Record cheetahs in Namibia
The biggest male cheetah ever recorded in all of Namibia weighed 64 kg and the smallest one weighed 31 kg.
The biggest female ever recorded in all of Namibia weighed 51 kg and the smallest female ever recorded weighed 26 kg.(4)


Total average in Namibia
For both genders cheetahs average 41kg in Namibia (n=759). In Namibia  cheetahs average 44kg for males (n=499) and 37 kg for females (n=260).(4)(3)(1)(16)


Botswana
Okavango delta
Cheetahs in Okavango delta,Botswana average 62 kg for males (n=7) .The biggest cheetah recorded in this sample and the biggest cheetah ever recorded in Botswana weighed 69 kg and was named Legolas.(12)


Kgalagadi and Ghanzi districts 
In this place males average 45 kg for males (n=24) and 39 kg for females (n=17).Average for both genders would be 43 kg.(5)


Total average in Botswana
In Botswana cheetahs average 49kg for males (n=31) and females average 39 kg for females (n=17). For both genders cheetahs average 45 kg (n=83) in Botswana.(5)(6)(12)



Kenya
Masai mara
Cheetahs in Masai Mara,Kenya average 32 kg for males (n=3) and 44 kg for females (n=8).Cheetahs for both genders average 41 kg in Masai mara (n=11).(5)


Unknown part of Kenya
In an unknown part of Kenya cheetahs average 61 kg for males (n=4) and 52 kg for females (n=2).For both genders cheetahs average 58 kg (n=6) in this area.The biggest male in this region of Kenya weighed 64kg and the smallest male in this region weighed 58 kg.The biggest female in this region weighed 63 kg and the smallest one weighed 41 kg.(1)


Total average in Kenya
Cheetahs in Kenya average 49 kg for males (n=7)n and and 44 kg for females (n=14).Cheetahs for both genders average 46 kg in kenya (n=21).(5)(1)


South Africa
Kalahari Gemsbok national park
 In Kalahari Gemsbok national park cheetahs weigh 54 kg for males (n=7) and 43 kg for females (n=6).Cheetahs for both genders average 47 kg in Kalahari Gemsbok national park(n=21).The biggest male ever recorded in Kalahari Gemsbok national park weighed 59 kg and the smallest male ever recorded in Kalahari Gemsbok national park weighed 39 kg.The biggest female ever recorded in Kalahari Gemsbok national park weighed 48 kg and the smallest female weighed 36 kg. (1)


Unknown area in South Africa
Cheetahs in an unknown area of South Africa Averaged 55kg for males (n=4) and 49 kg for females (n=2). For both genders cheetahs weigh 53 kg in this area (n=6).The biggest male in this area weighed 62 kg and the smallest male weighed 50 kg.The biggest female in this area weighed 58 kg and the smallest one weighed 39 kg.(1)


Record cheetahs in South Africa
The biggest male cheetah in South Africa weighed 62 kg and the smallest male ever recorded in this South Africa weighed  39 kg.The biggest female ever recorded in South Africa weighed 58 kg and the smallest female weighed 36 kg.(1)


Total average
Cheetahs in South Africa average 54 kg for males (n=11) and 45 kg for females (n=8).Cheetahs for both genders average 50 kg in South Africa (n=19).(1)


Tanzania
Serengeti
Cheetahs in Serengeti,Tanzania average 41 kg for males (n=23) and 36 kg for females (n=19).Cheetahs for both genders average 39 kg in the serengeti.The biggest male cheetah ever recorded in Serengeti weighed 51 kg and the smallest male ever recorded in Serengeti weighed 29 kg.The biggest female ever recorded in the serengeti weighed 43kg and the smallest female ever recorded in the serengeti weighed 21 kg.(1)


Zambia
The only male cheetah weighed in Zambia was male which weighed at 54 kg.(14)



Other wild cheetah weights


In the Namibian ranchland cheetahs of both averaged 37 kg (n=3).(15)



Cheetah Conservation Botswana 2014
Cheetah conservation Botswana 2014 gives 42 kg for males and females (n=57) in Serengeti and Masai Mara.(12)




Total average



Total average for wild male cheetahs:45 kg (n=700)


Total average for wild female cheetahs:38 kg (n=325)


Total average for both genders in wild:42 kg (n=1025)


Captive
Functional anatomy of the cheetah (Acinonyx jubatus) forelimb gives 33 kg average for 5 cheetahs.Males averaged 33 kg (n=3) and females averaged 38 kg (n=2).


Power output of skinned skeletal muscle fibres from the cheetah (Acinonyx jubatus) gives 41kg for one cheetah male.

Quasi-steady state aerodynamics of the cheetah tail gives 32 kg for 7 captive cheetahs.





Cheetah Husbandry Manual Volume III gives 44 kg for 82 male cheetahs





An acoustic analysis of purring in the cheetah (Acinonyx jubatus) and in the domestic cat (Felis catus) gives 68 kg for cheetah male named Caine.



Skeletal muscle histology and biochemistry of an elite sprinter, the African cheetah gives 47 kg for 3 cheetahs

EVALUATION OF A FIXED-DOSE COMBINATION OF BUTORPHANOLAZAPERONE-MEDETOMIDINE (BAM) FOR CHEMICAL IMMOBILISATION OF AFRICAN LION, BLESBOK, AND CHEETAH gives 54 kg for 6 male cheetahs and 48 kg for 6 female cheetah.







Total average for captive males:44 kg (n=97)





Total average for captive females:44 kg(n=9)





Total average for captive cheetahs:44 kg (n=106)







Semi-captive





Ultrasonographically determined renal values and comparisons to serum biochemistry renal variables in aged semi-captive cheetahs (Acinonyx jubatus) gives 40 kg for semi-captive male cheetahs (n=17) and 35 kg for semi-captive female cheetahs (n=10).As species cheetahs averaged 38 kg (n=27).The biggest male in this sample weighed 46 kg and the smallest male in this sample weighed 32 kg.The biggest female in this sample weighed 41 kg and the smallest female in this sample weighed 26 kg.





Africat

Semi-captive cheetahs weighed by africat averaged 42 kg for males (n=4) and 39 kg for females (n=2).As a species cheetahs averaged 41 kg (n=6).The biggest male in this sample weighed 47 kg and the smallest male weighed 37 kg.The biggest female weighed 42kg and the smallest female weighed 36 kg.




Benign Pigmented Dermal Basal Cell Tumor in a Namibian Cheetah (Acinonyx jubatus) gives 47 kg for one male cheetah.

Total average for semi captive cheetah males:40 kg (n=22)








Total average for semi-captive cheetah females:36 kg (n=12)





Total average for both genders (semi-captive):39 kg (n=34)







References:

1.Aspects of Cheetah (Acinonyx jubatus) Biology, Ecology and Conservation Strategies on Namibian Farmlands

2.Ultrasonographically determined renal values and comparisons to serum biochemistry renal variables in aged semi-captive cheetahs (Acinonyx jubatus) 

3.Africat Foundation

4.Morphology, Physical Condition, and Growth of the Cheetah (Acinonyx jubatus jubatus) 

5.Regional variation in the cheetah (Acinonyx jubatus) revisited: Morphology of wild and captive populations
6.Regional variation in body size of the cheetah (Acinonyx jubatus

7.Power output of skinned skeletal muscle fibres from the cheetah (Acinonyx jubatus)

8.Quasi-steady state aerodynamics of the cheetah tail

9.Cheetah Husbandry Manual Volume III

10.Functional anatomy of the cheetah (Acinonyx jubatus) forelimb 

11.Some Weights and Measurements of Large Mammals

12.Cheetah Conservation Botswana 2014-2015

13.Benign Pigmented Dermal Basal Cell Tumor in a Namibian Cheetah (Acinonyx jubatus


14.Arnoldia Rhodesia 1967


[b]15.Skeletal muscle histology and biochemistry of an elite sprinter, the African cheetah[/b]
[b]16.[/b]
[b][b]Spatial and Temporal Habitat Use by GPS Collared Male Cheetahs in Modified Bushland Habitat[/b][/b]
17.EVALUATION OF A FIXED-DOSE COMBINATION OF BUTORPHANOLAZAPERONE-MEDETOMIDINE (BAM) FOR CHEMICAL IMMOBILISATION OF AFRICAN LION, BLESBOK, AND CHEETAH 
18.Cheetahs (Acinonyx jubatus) running the gauntlet: an evaluation of translocations into free-range environments in Namibia







^UPDATED
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Canada Acinonyx sp. Offline
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#26

Credits to @TheNormalGuy for these weights.

Namibian farmlands
Spatial and Temporal Habitat Use by GPS Collared Male Cheetahs in Modified Bushland Habitat
https://html.scirp.org/file/_5-1620360_1.htm


Captivity:
EVALUATION OF A FIXED-DOSE COMBINATION OF BUTORPHANOLAZAPERONE-MEDETOMIDINE (BAM) FOR CHEMICAL IMMOBILISATION OF AFRICAN LION, BLESBOK, AND CHEETAH 

"ResultsGrinata from six male cheetahs weighing 54.1 ± 2.8 kg (range 50-57 kg) and from six female animals weighing 47.8 ± 5.6 kg (range 37-54 kg) were used in this study."


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https://dspace.emu.ee/xmlui/bitstream/handle/10492/6226/20210107_AS_PhD_thesis_final_10.11_Alo.pdf?sequence=4&isAllowed=y
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#27

Credits to @TheNormalGuy for these weights too

49 kg (n=4) for male cheetahs
40 kg (n=5) for female cheetahs
Source:Cheetahs (Acinonyx jubatus) running the gauntlet: an evaluation of translocations into free-range environments in Namibia


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#28

Dolph C. Volker Pers.comm.

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The captive cheetah named 'Gabe' was just under 45 kg or 100 pounds.
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Cheetah skull measurements and remarks on cheetah skull morphology in scientific literature

Cheetah Skull morphology



Skull length
Namibia
Namibian farmlands
Male cheetahs in the namibian farmland have a skull length of 23.4 cm (n=54) and females have a skull length of 21.8 cm (n=26).The smallest male had a skull length of 21 cm and largest 26.7 cm.The smallest female had a skull length of 18.5 cm and largest 24.1 cm.(1)


Otjiwarongo
Male cheetahs here had an average skull length of 20.3 cm (n=89) and females 19.1 cm (n=47).The largest male had a skull length of 23 cm and the largest female 21 cm.(3)


Etosha National Park
Male cheetahs here have an average skull length of 15.7 cm (n=298) and female cheetahs have an average skull length of 15.1 cm (n=146).(2)


Total average in Namibia
Male cheetahs had an average skull length of 17.5 cm (n=441) and female cheetahs 16.7 cm (n=191).(1,2,3)


Botswana
Okavango delta 
Male cheetahs in the Okavango delta have 22.7 cm (n=7).(5)


Ghanzi and Kgalagadi District
Male cheetahs in this area have an average skull length of 16.3 cm (n=24) and female cheetahs have an average skull length of 15.6 cm (n=17).(4)


Total average of Botswana
Male cheetahs in Botswana have an average skull length of 17.7 cm (n=31) and females have an average skull length of 15.6 cm (n=17). (4,5)


India
2 female cheetahs in India had an average skull length of 17.5 cm.(6)


Iraq
One asiatic cheetah in Iraq had a skull length of 17.2 cm(7)



Other skull lengths
Christiansen et al 2007 gives 14.9 cm for 9 cheetah skulls from museums of unknown genders and ages. Adolffsen et al 2005 gives 17.2 cm for one cheetah skull, gender and age unknown. Cheetah Conservation Botswana 2014-15 gives 15.6 cm for 57 cheetahs of both genders from Serengeti and Masai Mara. Anirudh et al gives a skull length of 15.9 cm.(5,9,10)



Skull width


Namibia
Namibian farmlands
Male cheetahs in the namibian farmlands have an average skull width of 14.6 cm (n=84) and females have an average skull width of 13.3 cm (n=37).The largest male had a skull width of 17.2 cm and the smallest male had a skull width of 12.4 cm.The largest female had a skull width of 14.7 cm and the smallest female had a skull width of 10.8 cm.(1)


Otjiwarongo District
Males in the Otjiwarongo District have an average skull length of 15.88 cm (n=89) and females have an average skull width of 14.93 cm (n=47).The largest male cheetah had a skull width of 18.50 cm and the largest male cheetah had a skull width of 16 cm.(3)


Etosha National Park
Male cheetahs in Etosha national park have an average skull width of 14.1 cm (n=298) and female cheetahs have an average skull width of 13.2 cm (n=146).(2)


Total average in Namibia
Male cheetahs have an average skull width of 14.5 cm (n=470) in all of Namibia and female cheetahs have an average skull width of 13.5 cm (n=230).(1,2,3)


Botswana
Okavango delta
Male cheetahs in the Okavango delta have an average skull width of 15.4 cm (n=7).(5)


Kgalagadi and Ghazni Districts
Male cheetahs in these districts have an average skull width of 14.3 cm (N=24) and female cheetahs have an average skull width of 13 cm (n=17).(4)


Total average in Botswana
Male cheetahs have an average skull width of 14.5 cm (n=31) and female cheetahs have an average skull width of 13 cm (n=17) in all of Botswana.(4,5)


Canine length
Namibian farmlands
Male cheetahs in the namibian farmlands have an average upper canine length of 2.2 cm (n=87) and females have an average upper canine length of 1.6 cm (n=37). Male cheetahs in the namibian farmlands have an average lower canine length of 1.6 cm (n=86) and females 1.4 cm (n=35).The largest male cheetah ever recorded had a upper canine length of 2.6 cm and the smallest cheetah ever recorded had an upper canine length of 1.6 cm in the namibian farmlands.The largest female cheetah had an upper canine length of 2.3 cm and the smallest female cheetah ever recorded had an upper canine length of 1.7 cm in the namibian farmlands.The largest male cheetah ever recorded had a lower canine length of 2.1 cm and the smallest male cheetah had a lower canine length of 1 cm in the namibian farmlands.The largest female cheetah had a lower canine length of 2 cm and the smallest female cheetah had a lower canine length of 1 cm in the namibian farmlands.(1)


Otjiwarongo district
Male cheetahs have an average upper canine length of 2.3 cm (n=89) and female cheetahs have an average upper canine length of 2.2 cm (n=47) in the Otjiwarongo district. Male cheetahs have an average lower canine length of 1.7 cm (n=89) and female cheetahs have an average lower canine length of 1.5 (n=47) in the Otjiwarongo district.The largest male cheetah ever recorded had an upper canine length of 2.6 cm and a lower canine length of 2 cm in the Otjiwarongo district.The largest female cheetah ever recorded had an upper canine length of 2.2 cm and a lower canine length of 1.5 cm in the Otjiwarongo district.(3)


Etosha National Park
Male cheetahs have an average upper canine length of 2.1 cm (n=298) and female cheetahs have an average upper canine length of 2 cm (n=146) in the Etosha National Park. Male cheetahs have an average lower canine length of 1.6 cm (n=298) and female cheetahs have an average lower canine length of 1.4 cm (n=146).(2)


Total average in Namibia
Male cheetahs have an average upper canine length of 2.15 cm (n=474) and lower canine length of 1.6 cm (n=473). Female cheetahs have an average upper canine length of 1.56 cm (n=234) and an average lower canine length of  1.4 cm (n=321).(1,2,3)


Botswana
Kgalagadi and Ghazni districts
Male cheetahs have an average upper canine length of 2.3 cm (n=24) and an average lower canine length of 1.7 cm (n=24) in the Kgalagadi and ghanzi district. Female cheetahs have an average upper canine length of 2.1 cm (n=17)  and an average lower canine length of 1.5 cm (n=17).In the kgalagadi and ghazni districts.(4)


Other canine data
Meachen Samuels reports an average canine ML diameter of 8.82 mm and a canine crown height of 18.63 mm. BVM Anirudh reports a canine length of 30.13 mm and a canine crown height of 14.80 mm. Christiansen reports a upper canine length of 2.8 cm and a lower canine length of 2.08 cm for one cheetah specimen.(15,16)


Other Skull and Skull related Measurements
2 asiatic female cheetahs had a Condylobasal Length of 15.55 cm, Zygomatic width of 11.9 cm, Postorbital width of 5.2 cm, Interorbital width of 3.8 cm, Maxillary width of 4.8 cm, Mandible length of 11.9 cm, skull length-width ratio of 1.44 cm and a Total score of the skull of 29.05 cm. Christiansen reports a cheetah specimen that had a basicranial length of 10.7 cm, snout length of 6.46 cm, Jaw Joint to Carnassial length of 7.47 cm, Jaw Joint to canine length of 10.91 cm, lower jaw length of 12.95 cm and a gape of 6.68 cm. Christensen also reports a cheetah specimen to have a upper canine anteroposterior diameter of 2.82 cm, a upper canine lateromedial diameter of 0.89 cm, a lower canine anteroposterior diameter of 1.05 cm and a lower canine lateromedial diameter of 0.76 cm.Therrien reports a mandibular length of 11.84 cm for cheetahs.One asiatic cheetah from iraq had a condylobasal length of 15.63 cm, zygomatic breadth of 11.41 cm, interorbital breadth of 4 cm, braincase width of 6 cm, canine to molar distance if 5.32 cm and a mandible length of 11.96 cm. A cheetah from El Malfa, Egypt had a zygomatic breadth of 10 cm, canine to molar distance of 4.73 cm and a mandible length of 10.42 cm.In libya 2 female cheetahs had a condylobasal length of 13.44 cm. In Libya 3 female cheetahs had a zygomatic width of 10.06 cm, upper tooth row length of 4.7 cm and a mandible length of 10.28 cm.(6,7,8,10,17)



Skull structure
Cheetah skulls are highly domed in lateral aspect.The zygomatic arch is broad anteriorly and much narrower posteriorly.The Maxillary process is short and very little salient.The maxillary process does not in anyway overhand the infraorbital foramen.In postorbital processes tare very short and blunt.The vertical branch of the premaxilla protrudes only a short distance between the maxilla and the nasal.The nares are very large.The braincase is very rounded  and there is usually little evidence of a Sagittal crest except at the extreme posterior.Ventrally the palate is short and broadly triangular. The post dental palate has sharp sloping sides.The hamulars are broad hooks.The bullae is inflated but it is relatively small for the overall size of the skull.The mandible is not particularly strongly built.The dentition is distinctive.The cheek teeth are nominally 3.1 / 2.1. The space between the canine and the first premolar is usually narrow.The jaws have little sign of any postcanine gap.The canines exhibit little or no sign of furrowing.The miracinonyx species also have similar skull structure. Cheetah canines are small, hooked backwards and have grooves that are absent or faint. Nasals are Straight to Concave.(11,12,13)


Size of skull in comparison to the members of the puma lineage
Cheetahs have larger skulls than jaguarundis.There are no significant difference between pumas and cheetahs in the main axis of the skull (length, width,height) and 12 out of 20 skull measurements. Pumas score better in mastoid breadth,temporal fossa length, tooth row length and jaw width. Cheetahs score better in masseteric scar length, post orbital constriction breadth, condyle to length of jaw and orbit to premaxillae length.(14)



Total average
Average skull measurements for both genders
Skull length:17.08 cm (n=750)
Skull width:14.15 cm (n=748)
Upper Canine Length:2.08 cm (n=750)
Lower Canine Length:1.86 cm (n=836)
Condylobasal length:14.72 cm (n=5)
Zygomatic width:11.05 cm (n=5)
Postorbital Width:5.2 cm (n=2)
Interorbital width:3.86 cm (n=3)
Maxillary width:4.8 cm (n=2)
Mandible length:11 cm (n=7)
Skull Length Width Ratio=1.44 (n=2)
Total score of the skull:29.05 cm (n=2)
Basicranial length:10.7 cm (n=1)
Snout Length:6.46 cm (n=1)
Gape:6.68 cm(n=1)
Lower Jaw Length:12.95 cm (n=1)
Jaw Joint to Carnassial Length:7.47 cm (n=1)
Jaw Joint to Canine Length:10.91 cm (n=1)
upper canine anteroposterior diameter:2.82 cm (n=1)
 upper canine lateromedial diameter of 0.89 cm (n=1)
lower canine anteroposterior diameter:1.05 cm (n=1)
lower canine lateromedial diameter: 0.76 cm (n=1)
Braincase Width:6 cm (n=1)
Canine to Molar Distance:5.02 cm (n=2)
Upper Tooth Row Length:4.7 cm (n=3)


References
1.Morphology, Physical Condition, and Growth of the Cheetah (Acinonyx jubatus jubatus)
2.Regional variation in the cheetah (Acinonyx jubatus) revisited: Morphology of wild and captive populations
3.Africat
4.Regional variation in body size of the cheetah (Acinonyx jubatus
5.Cheetah Conservation Botswana 2014-15
6.Fauna of British India
7.The Cheetah (Acinonyx Jubatus 1776) in Egypt (Felidae, Acinonychinae)
8.Libyan Mammals:Cats of Libya
9.Comparative Biteforces and Canine Bending Strength in Feline and Sabertooth felids:implications for predatory ecology
10.Bite Forces, Canine Strength and Skull Allometry in Carnivores
11.Carnivores of West Africa
12.Wild Cats of the World, Sunquist and Sunquist 2002
13.Cranial Morphology of Five Felids
14.Skull morphology and functionality of extant Felidae (Mammalia: Carnivora): A phylogenetic and evolutionary perspective
15.Morphological convergence of the prey-killing arsenal of sabertooth predators
16.Comparing the dimensions of canine tooth in different species
17.Mandibular force profiles of extant carnivorans and implications for the feeding behaviour of extinct predators
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#30

Good job collecting this info. Are you sure the skulls from the Namibian farmlands were dried skulls? They seem to be very different from other Namibian cheetah skulls in size.
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Welcome to WILDFACT forum, a website that focuses on sharing the joy that wildlife has on offer. We welcome all wildlife lovers to join us in sharing that joy. As a member you can share your research, knowledge and experience on animals with the community.
wildfact.com is intended to serve as an online resource for wildlife lovers of all skill levels from beginners to professionals and from all fields that belong to wildlife anyhow. Our focus area is wild animals from all over world. Content generated here will help showcase the work of wildlife experts and lovers to the world. We believe by the help of your informative article and content we will succeed to educate the world, how these beautiful animals are important to survival of all man kind.
Many thanks for visiting wildfact.com. We hope you will keep visiting wildfact regularly and will refer other members who have passion for wildlife.

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