There is a world somewhere between reality and fiction. Although ignored by many, it is very real and so are those living in it. This forum is about the natural world. Here, wild animals will be heard and respected. The forum offers a glimpse into an unknown world as well as a room with a view on the present and the future. Anyone able to speak on behalf of those living in the emerald forest and the deep blue sea is invited to join.
I agree that there is probably a cycle in these things. We had a period of large powerful coalitions a while ago which thinned out the amount of rival coalitions. As a result a lot of cubs are currently making it to adulthood which is increasing the amount of coalitions in the reserve until there are too many.
I don't think the male lions care much about the exact number of sexual partners, defeated rivals or leaving a genetic legacy. They are just trying to make the best out of life.
It wouldn't be fair to compare a male lion that prefers to stick to one pride and defend it so well he raises multiple generations of cubs. With one that likes to take over prides and does so very well. But loses them quickly once he leaves to take another.
Both can be argued to be the most successful. The first for having a good long life as a pride male and most offspring. The second for having the most opportunities to mate and successful takeovers.
Perhaps the fairest comparison would be who lost the least fights as that is what each lion tries to avoid. But I don't think there are records on that.